SPHODRISTOCARABUS Géhin, 1885; type species Carabus adamsi Adams
species: 16
1.adamsi Adams, 1817 C,E-Caucasus; S-Azerbaydzhan; N-Iran
ssp. adamsi Adams, 1817 C,E-Caucasus:Terek River, Vladikavkaz env.- Telavi env.
=chiragricus Fischer, 1828 "thermal springs of Caucasus"(=Pyatigorsk, Kislovodsk env.)
=eichwaldi Fischer, 1828 "Kaukasus"
=chrysitis Motschulsky, 1839 "Alp.Cauc.:Tuschetia"(LT)
=thomsonianus Géhin, 1876
=subcyaneus Kraatz, 1878, nec Illiger, 1802; nbm C-Caucasus:Kazbegi env.; "?Caucasus"
=porphyrobaphes Ganglbauer, 1887 ?LNF(?E of Vladikavkaz)
=adamsianus Mandl, 1975 "Caucasus"
=svaneticola Deuve, 2002 "Svanetia"
~kabardaensis Deuve, 2002, nec kabardensis Kurnakov, 1970 "Kabarda"
=pseudojanthinus Deuve, 2002 LNF(unlabelled HT)
ssp. hollbergi Mannerheim, 1827 E-Cauc.:Shakh D., Kurush; Azerbaydzhan:Kirovabad, Khanlar, Karabakh Mts.; "Dagestan"(LT)
=separandus Kraatz, 1881 "Elisabethpol"(=Kirovabad; =Gyandzha)
~eichwaldianus Géhin, 1885; ?in litt.
~grusianus Roeschke; in litt.
~separandulus Mandl, 1975; un LNF(unlabelled LT)
=adjikentensis Deuve, 2002 Azerbaydzhan:"Adjikent"(=Adjikend; ca.15km SE Gyandzha); 1m
=khanlarensis Deuve, 2002 Azerbaydzhan:"Helenendorf"(=Khanlar; 9km S Gyandzha)
=tchaikentensis Deuve, 2002 Azerbaydzhan:"Tchaikent"(=Chaykend, ca.17km SE Gyandzha); 1m
ssp. serriae Deuve, 2002 N-Iran:Azerbaydzhan:E-part:Kaleybar env.
ssp. separatus Lapouge, 1907 SE-Azerbaydzhan:Lenkoran; N-Iran:Gilan:Asalom, Masuleh; "Talysh"
ssp. persa Lapouge, 1924 N-Iran:Elborz Mt.R.:Ramsar env. to Sari env.; "W-Elborz"(HT lost); Kelardasht (NT)
~longicollis Mandl, 1955, nec Duftschmid, 1812 "Persia, Elburs"
~sahelae Lassalle, 2002; un N-Iran:Mazandaran:betw.Behshahr/Gorgan:Mts. S of Galugah, 1800m
ssp. korgei Heinz, 1970 N-Iran:Mazenderan:Khalus valley:pass N of Gakhsar(~36:10N/51:19E)
=delirensis Deuve, 2000 N-Iran:Mazendaran:SW Chalus:Delir, 36:21N/51:08E
ssp. haerii Heinz, 1973 N-Iran:Gilan:E of Rudbar:Dare-dasht
=larkhaniensis Deuve, 2001 N-Iran:Gilan:Espeyli env.:Larkhani Mts., 1400 m; betw.Siyahkal/Espeyli; Balarud env.
ssp. bararii Deuve, 2000 N-Iran:Mazendaran:Ramsar env.:Ney Dasht, 300-350m
ssp. samaorum Rapuzzi, 2005 NE-Iran:Golestan:Gorgan env.(20km S Gorgan:Nahar Horan)
2.biroi Csiki, 1927; nom.pro kindermanni Chaudoir NC-Turkey
ssp. biroi Ménétriés, 1832 NC-Turkey:Samsun:Kavak env.; Amasya
~kindermanni Chaudoir, 1850, nec Waltl, 1838 Amasya
ssp. iorbi Deuve & Charet, 1998 NC-Turkey:Samsun:W of Ladik
ssp. leonardoi Deuve, 2005 NC-Turkey:Sinop:30km E Ayancik; 1f
3.bohemani Ménétriés, 1832 Azerbaydzhan/Iran borders region; N-Iran:Gilan, Mazandaran
ssp. bohemani Ménétriés, 1832 Azerbaydzhan/Iran borders region:Lenkoran, Ardabil env.; "Mts.of Talysh"
=yvdouarini Deuve & Doline, 2003 Azerbaydzhan:Lenkoran anv.
Hirkansky Les, Astara, Masilli)
ssp. esperanzae Heinz, 1970 N-Iran:Gilan:NW of Safid valley:Masuleh, Asalam
ssp. esfandiarii Heinz, 1973 N-Iran:Mazandaran:Sardab-rud valley:Rudbarak, S of Chalus:Kelardasht
ssp. morvani Morvan, 1976 N-Iran:Mazendaran:Elborz Mt.R.:Chorteh, 1800m(=SW of Ramsar)
ssp. rosicolor Deuve, 2001 N-Iran:Gilan:Espeyli env.:Larkhani Mts., 1400 m
4.coruhnehriensis Cavazzuti, 1990 NE-Turkey:Artvin:Coruh valley:25-30km W Yusufeli; ?Rize:Ovit Pass:N-slope
5.elegantulus Motschulsky, 1850 N-Iran:E-Elborz Mt.R.; Iran/Turkmenistan:Kopet Dag Mt.R.
ssp. elegantulus Motschulsky, 1850 N-Iran:E-Elborz:Gorgan env.(="Astrabad"), Bojnurd; Iran/Turkmenistan:Kopet Dag Mt.R.
=astrabadensis Semenov, 1903 "Mts. of Astrabad Province"(=Gorgan)
=parthicus Lapouge, 1924 "Budschnurd Mts."(=Bojnurd env.)
ssp. sarafrazii Deuve, 2000 N-Iran:Mazandaran:Sari:Kiyasar:Vesmine Mts.(Kiyasar:36:14N/53:22E)
ssp. dinae Rapuzzi, 2005 NE-Iran:Golestan:S of Gaz, 36:39N/53:58E
6.enigmaticus Heinz, 1980 NE-Turkey:Erzurum, Artvin, Kars
ssp. enigmaticus Heinz, 1980 NE-Turkey:Erzurum:Ispir env.(10km NW Ispir, 1500-1700m)
ssp. reiseri Heinz, 1980 NE-Turkey:Artvin:NW of Yusufeli:Yaylalar, Altiparmak env.
ssp. exiliatus Cavazzuti, 1990 NE-Turkey:Artvin:Kiliçkaya env.
eve Dag Mts.:1800-2000m
ssp. citamgineus Lassalle & Charet, 1992 NE-Turkey:Kars:Sarikamis, Göle
=bechteli Charet, 1996 Kars:Göle
7.georgia Cavazzuti, 1984 CE-Turkey:Tunceli env.(10km NE of Tunceli)
=hardei Heinz, 1990 CE-Turkey:Tunceli:Kötek-Yayla, Gozen env.
8.giachinoi Cavazzuti, 1991 NE-Turkey:Gümüshane:NE of Siran:Tersundagi Pass, 1800m; 4f
9.gilnickii Deyrolle, 1869 NE-Turkey:Trabzon, Rize, Giresun, Erzurum
ssp. gilnickii Deyrolle, 1869 NE-Turkey:Trabzon env.:Macka, Zigana Pass, Soganli Pass, Ovit Pass
=giulielli Thomson, 1875
=acutus Lapouge, 1904 Trabzon:"Kolat Dag"(=Mts.nr.Zigana Pass)
~viridifossulatipennis Breuning, 1964; un Trabzon:Balabandag W of Macka
~brignolii Breuning & Ruspoli, 1970; un Trabzon:"Kolat Dag"(=Mts.nr.Zigana Pass)
=yakalamakus Lassalle, 1993 NE-Turkey:Giresun:S of Dogankent River:steppes W of Kürtün
=ovoideus Machard, 1994 Trabzon:Soganli Pass S of Caykara
=pseudoelongatus Machard, 1994 Rize:betw.Sivrikaya/Ovit Pass
=lavezorum Darge, 1994 Rize:Ovit Pass:N-slope
ssp. hasandaghensis Darge & Basquin, 1990 NE-Turkey:Erzurum:N of Ispir:Hasan Dag
ssp. brunneicupreus Deuve, 1994 NE-Turkey:Rize:Çat
10.heinzi Breuning, 1964 NE-Turkey:Giresun, Ordu
ssp. heinzi Breuning, 1964 NE-Turkey:Giresun:Egribel Pass, Kulakkaya, Tamdere env.
ssp. mateuellus Deuve, 2003 NE-Turkey:Ordu:Gölköy env.(pass betw.Mesudiye/Gölköy)
~matteui Breuning & Ruspoli, 1970; un NE-Turkey:Ordu:Gölköy env.(pass betw.Mesudiye/Gölköy)
11.kurdicus Heinz, 1975 CE,SE-Turkey:Bitlis, Bingöl, Siirt
ssp. kurdicus Heinz, 1975 CE-Turkey:Bitlis:Tatvan, Resadiye env.
~schuberti Breuning, 1970, nec Breuning, 1968 Bitlis:Tatvan env.
=basquini Darge, 1990 Bitlis:Resadiye/Gevas:Alacabük Dag, 1770m
ssp. machardianus Machard, 1986 CE-Turkey:Bingöl env.; 2f
ssp. kurdaorum Lassalle, 1992 SE-Turkey:Siirt:Yolbilen
12.macrogonus Chaudoir, 1847 N-Turkey:Adapazari to Artvin
ssp. macrogonus Chaudoir, 1847 NE-Turkey:Trabzon env., Macka valley, Zigana Pass; Giresun; LNF
=theophilei Deyrolle, 1869 Trabzon env.
=kolatensis Lapouge, 1924 "Kolat Dagh nr.Trabzon"(=Mts.nr.Zigana Pass)
=koselerensis Machard, 1991 Giresun:betw.Tirebolu/Köseler
=giresunensis Deuve, 1992 "Anatolia, Giresun"(exactly:?Balaban Dag=Mts.SW of Yavuzkemal)
=lemeslei Machard, 1995 Trabzon:S of Caykara:Köknar, 1800m
ssp. pseudokolatensis Heinz & Korge, 1967 NE-Turkey:Artvin:Kackar Dag:N-slope; Rize:Ilica, Ardesen valley, Ikizdere
=rizensis Ledoux, 1975 Rize:S of Pazar
=michaelae Darge, 1994 Rize:Ovit Pass:N-slope
ssp. azaleae Cavazzuti, 1986 NE-Turkey:Ordu:40km SE Ordu:Cambaşi(=E of Topcam)
ssp. resplendens Breuning, 1934 NC-Turkey:Amasya:Borabay; Samsun:Ayvacik, Carsamba, Ladik; Ordu:Ünye
=chambruni Breuning & Ruspoli, 1970 Samsun:Akpinar nr.Carsamba
=cavazzutianus Casale, 1973 Ordu:S of Ünye
=zanderi Battoni & Blumenthal, 1973 Samsun:Ladik
~eiselti Mandl, 1975; un Ordu:betw.Akkus/Ünye
ssp. akkusensis Breuning & Ruspoli, 1970 NE-Turkey:Ordu:Akkus, Gulebi, Ulubey env.; Tokat:Niksar
~akkusensis Breuning, 1964; un Ordu:Akkus
~ingloriosus Breuning & Ruspoli, 1970; un Ordu:Akkus
=guergentepeensis Blumenthal, 1975 Ordu:betw.Gölköy/Ulubey:Gurgentepe Pass
=lilianae Casale & Cavazzuti, 1976 Ordu:Gölköy
~daumani Breuning & Ruspoli, 1976; un Ordu:Ulubey
ssp. simonianus Breuning & Ruspoli, 1970 NE-Turkey:Ordu:betw.Mesudiye/Koyulhisar; Gölköy, Mahmudiye Sivas:Koyulhisar
~pseudosimonianus Mandl, 1975; un Ordu:Gölköy
ssp. diogenei Cavazzuti, 1991 NC-Turkey:Sinop:Gokcebel, Incemeydan, Hatip, Buyukduz, Turkeli
ssp. schweigerinae Schweiger, 1962 NW-Turkey:Bolu(Abant Lake)
=goettingi Schweiger, 1964 Bolu:Abantayagi
=pseudoschweigerinae Schweiger, 1964 Bolu:Abantayagi
ssp. drannadaghensis Breuning, 1964 NW-Turkey:Kastamonu:Ilgaz, Yaraligöz; Sinop:Dranaz Dag, Cangal Dag
=clientulus Schweiger, 1969 Istanbul City (?introduced)
~mollioni Breuning & Ruspoli, 1976; un Sinop:Cangal Dag 40 km W Sinop:S of Ayancik
ssp. keltepensis Vigna-Taglianti & Bruschi, 1976 NW-Turkey:Adapazari:Sapanca env.; Kocaeli:Goek Dag nr.Izmit
13.pavesii Cavazzuti, 1992 W-Turkey:Izmir:Bornova; 1m; probably introduced macrogonus-specimen
14.scovitzi Faldermann, 1835; cos SW-Georgia:Batumi, Makharadze env; NE-Turkey:Artvin, Rize
ssp. scovitzi Faldermann, 1835; cos SW-Georgia:Batumi, Makharadze env.; LNF(?Batumi env.)
~sovitzi Faldermann, 1835; ?iss
ssp. otingoensis Deuve, 1993 NE-Turkey:"Artvin:Karckal-Otingo"(=Karcal Dagi:Balci env.)
ssp. hedini Heinz, 1973 NE-Turkey:Rize & Artvin:Ayder, Yaylalar env.(Ardesen valley:Kavran-yayla)
ssp. bouilloni Breuning, 1976 NE-Turkey:Artvin:Murgul nr.Borçka
ssp. czipkai Heinz, 1980 NE-Turkey:Artvin:Artvin env.(forests above Artvin, 1600-1800m)
ssp. sarigoelensis Deuve & Charet, 1992 NE-Turkey:Artvin:35km W Sarigol
15.tokatensis Roeschke, 1898 NC,CE-Turkey:Tokat, Gümushane, Tunceli, Erzurum
ssp. tokatensis Roeschke, 1898 NC-Turkey:W of Tokat:Topcam Dag Massif; Tokat
ssp. almushensis Mandl, 1975 NC-Turkey:E of Tokat:Almus env.; Yildiz Daglari; Sivas:Koyulhisar
ssp. kosensis Cavazzuti, 1984CE-Turkey:Gümüshane:Köse(pass N of Köse), Siran, Kelkit, Torul env.
ssp. kirikensis Staven, 1995 CE-Turkey:Erzurum:W of Kirik, SE of Kirik
ssp. simonae Cavazzuti, 1984 CE-Turkey:Tunceli:Munzur Daglari:Pülümür, Ovacik
ssp. karasudominus Cavazzuti, 1991 CE-Turkey:Erzurum:Karasu Daglari:Cat env.; 2f
16.varians Fischer, 1827 Armenia, Georgia; SW,SC-Caucasus; NE-Turkey
ssp. varians Fischer, 1827 Armenia:Sevan, Dilidzhan, Yerevan; Georgia:Tbilisi, Somkhet Mt.R.; "Caucaso cisalpino"
=sieversi Ganglbauer, 1887 Caucasus Minor:Trialetsky Mt.R., Bakuriani; "Manglis nr.Tiflis"(=Tbilisi env.)
~praevarians Mandl, 1975; un "Somchetia"
~somcheticus Mandl, 1975; un Armenia:Dilidzhan
ssp. armeniacus Mannerheim, 1830 NE-Turkey:Erzurum(Kop Dag S of Trabzon); Artvin; Georgia:Caucasus Minor, C-Caucasus
=incatenatus Mannerheim, 1830 Kop Dag S of Trabzon
~confusus Fischer, in litt.
=fulminans Kraatz, 1878 ?Borzhomi
=subincatenatus Kraatz, 1878 ?Borzhomi
=repletus Reitter, 1884 SC-Caucasus:upper Inguri & Rioni River; "Svanetia"
~mutabilis Géhin, 1885; un
=korbianus Ganglbauer, 1887 S-Georgia:Batumi env.; NE-Turkey:Artvin:Hopa
=laevilineatus Ganglbauer, 1887 SC-Caucasus:Kodorsky Mt.R., Egrissky Mt.R., Tkvarcheli, Zugdidi env.
=decoloratus Reitter, 1896 ?"Svanetia"
~pseudoarmeniacus Breuning, 1935; un S-Georgia:"Achalzich"(=Akhaltsikhe)
~voriseki Novotný, 1975; un ?Trialetsky Mt.R.:Mt.Ckhra-Ckkhara
=arsiyandagensis Machard, 1985 Kars:NW of Hanak:Arsiyan Dag
=sarikamisensis Cavazzuti, 1986 Kars:Sarikamis
=robichei Auvray, 1994 Artvin:Zeytinlik
ssp. scintillus Reitter, 1884 C-Caucasus:S-slope:"Svanetia"(alpine form)
ssp. janthinus Ganglbauer, 1887, nec Duftschmid, 1812; nbm W-Caucasus:Gagrinsky Mt.R.-Tuapse; Maykop env.; Slavyansk:Krasny Les; Mt.Achishkho(LT)
~ciscaucasicus Mandl, 1975; un Dagomys
~dvorshaki Mandl, 1975; un SW-Caucasus:Pitsunda env., Adler; Bzybsky Mt.R.
~manfredschmidi Mandl, 1975; un Maykop env:Suntuk
ssp. rugatus Breuning, 1934 W-Caucasus:Sukhumi, Novy Afon, Tsebelda, Amtkel, Atsgara, Abkhazsky Mt.R.(Marukh Pass)
~novotnyorum Mandl, 1975; un Sukhumi
ssp. alagoensoides Mandl, 1975 W-Armenia:Mt.Aragats; "Araxestal"(=Araks valley)
~alagoensis Reitter, in litt.
~alagoensoides Mandl, 1955; un "Araxestal"(=Araks valley)
ssp. blumenthaliensis Heinz & Korge, 1967 NE-Turkey:Rize:Kackar Dagi:N-slope(Ilica, Ardesen valley)
ssp. basilewskianus Breuning & Ruspoli, 1970 Artvin:Hopa env.(Ciftekoprüler betw.Hopa/Borcka); ?Ogdem env.
=aabadensis Lassalle, 1991 NE-Turkey:Artvin:Kafkasor nr.Artvin, 1800m
ssp. aequicostalis Staven & Heinz, 1995 NE-Turkey:NE of Artvin:Dalis Dag
ssp. rotundithorax Deuve, 2003; nom. pro rotundicollis Mandl NE-Turkey:Artvin:Karçal Dagi Massif; ?Tunceli; Kartsch Chal(=Karçal Dagi)
~rotundicollis Mandl, 1955, nec Marsham, 1802 Kartsch Chal(=Karçal Dagi)
Carabus (Sphodristocarabus) varians Fischer, 1827
